Campbell Biology 12th Edition, AP® Edition © 2021 

Campbell Biology AP® Edition, published by Pearson, delivers trusted, accurate, and current instruction that builds upon the tools, visuals, and activities Neil Campbell used to encourage participation and engage students.

  • Prepares students for AP Course and Exam success.
  • Ties together key themes and big ideas
  • Strategic integration of texts and artwork
  • This program is accompanied by Pearson’s Mastering® platform with eText

AP® Biology Program with Digital Integration

Campbell Biology AP® Edition upholds the Campbell tradition of accuracy, currency, and passion for teaching and learning AP Biology.


Support for AP

Offers AP-specific chapter level features that clearly correlate the textbook’s content to the AP Curriculum Framework and prepare students for AP Exam success.


Relevant Instruction

Students work on real-world problems that use easy entry points to build investment and engagement with their learning.


Innovative Pedagogy

Scientific Skills Exercises in every chapter use real data to build key skills needed for biology, including data analysis, graphing, experimental design, and math skills.



The NEW Pearson LabBench features 13 investigations that reinforce key concepts from the lab outside the laboratory and prepare students for success on the AP exam and beyond.

AP® Biology Program Solution

  • Develop a Conceptual Framework
  • Make Connections to Biology
  • Build Key Scientific Skills
  • Instruction for Modern Learners
  • Visualize Key Biological Concepts
  • Chapter Openers
    Chapter Openers introduce each chapter and feature a question answered with a clear, simple image to help students visualize and remember concepts as they move through the chapter.
  • Study Tips
    Each opener includes a study tip and points students to interactive media and study tools in Mastering Biology.
  • The Big Picture

    Show the “big picture” relationships between content from different chapters.

  • Connections Tutorials

    Connections Tutorials are assignable and automatically graded in Mastering Biology and include answer-specific feedback for students.

  • Scientific Skills Exercises
    Scientific Skills Exercises in every chapter of the text use real data to build key skills needed for biology.
  • Develop Skill Sets
    Covering data analysis, graphing, experimental design, and math skills.
  • Pre-Lab Materials
    New Pre-lab materials challenge students to pose insightful questions and think critically before they conduct an experiment.
  • Post-Experiment Evaluation
    After the experiment, they evaluate their data, apply mathematical techniques, and compare the results against their expected outcomes. 
  • Howard Hughes Medical Institute Movies
    Documentary-quality movies from the Howard Hughes Medical Institute delve into biology topics such as the double helix and evolution.
  • BioFlix® 3D Animations
    BioFlix® 3D animations visualize complex ideas and can be presented in class, via the Study Area or eText, or accessed through the Mastering platform.

Take a Deeper Look at Campbell Biology AP® Edition



Get to know Campbell Biology AP® Edition with Mastering® Biology with Pearson eText.

Online Platform for your AP® Biology Course



The Pearson Mastering® platform delivers engaging, dynamic learning opportunities—focused on course objectives and responsive to reach student’s progress—proven to help students absorb course material and understand difficult concepts.

AP Test Prep from Pearson

  • Support for Students

Test Prep Workbook

The Test Prep workbook authored by award-winning AP® Biology instructors Fred and Theresa Holtzclaw helps students develop science skills and practice for the AP exam. Available for separate purchase.

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Correlation for AP®

Our solutions for AP® are designed to support and correlate the College Board's Course and Exam Descriptions for each corresponding course.

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More About Campbell Biology

  • Lisa A. Urry Author Bio
    Lisa A. Urry (Chapter 1 and Units 1-3) is Professor of Biology at Mills College. After earning a B.A. at Tufts University, she completed her Ph.D. at the Massachusetts Institute of Technology (MIT). Lisa has conducted research on gene expression during embryonic and larval development in sea urchins. Deeply committed to promoting opportunities in science for women and underrepresented minorities, she has taught courses ranging from introductory and developmental biology to an immersive course on the U.S./Mexico border. 
  • Michael L. Cain Author Bio

    Michael L. Cain (Units 4, 5, and 8) is an ecologist and evolutionary biologist who is now writing full-time. Michael earned an A.B. from Bowdoin College, an M.Sc. from Brown University, and a Ph.D. from Cornell University. As a faculty member at New Mexico State University, he taught introductory biology, ecology, evolution, botany, and conservation biology. Michael is the author of dozens of scientific papers on topics that include foraging behavior in insects and plants, long-distance seed dispersal, and speciation in crickets. He is also a coauthor of an ecology textbook.

  • Steven A. Wasserman Author Bio
    Steven A. Wasserman (Unit 7) is Professor of Biology at the University of California, San Diego (UCSD). He earned an A.B. from Harvard University and a Ph.D. from MIT. Working on the fruit fly Drosophila, Steve has done research on developmental biology, reproduction, and immunity. Having taught genetics, development, and physiology to undergraduate, graduate, and medical students, he now focuses on introductory biology, for which he has been honored with UCSD’s Distinguished Teaching Award.
  • Peter V. Minorsky Author Bio
    Peter V. Minorsky (Unit 6) is Professor of Biology at Mercy College in New York, where he teaches introductory biology, ecology, and botany. He received his A.B. from Vassar College and his Ph.D. from Cornell University. Peter taught at Kenyon College, Union COllege, Western Connecticut State University, and Vassar College; he is also the science writer for the journal Plant Physiology. His research interests concern how plants sense environmental change. Peter received the 2008 Award for Teaching Excellence at Mercy College.
  • Rebecca B. Orr Author Bio
    Rebecca B. Orr (Ready-to-Go Teaching Modules, Interactive Visual Activities, eText Media Integration) is Professor of Biology at Collin College in Plano, Texas, where she teaches introductory biology. She earned her B.S. from Texas A&M University and her Ph.D. from University of Texas Southwestern Medical Center at Dallas. Rebecca has a passion for investigating strategies that result in more effective learning and retention, and she is a certified Team-Based Learning Collaborative Trainer Consultant. She enjoys focusing on the creation of learning opportunities that both engage and challenge students.
  • Neil A. Campbell Author Bio
    Neil A. Campbell (1946-2004) earned his M.A. from the University of California, Los Angeles and his Ph.D. from the University of California, Riverside. His research focused on desert and coastal plants. Neil’s 30 years of teaching included introductory biology courses at Cornell University, Pomona College, and San Bernardino Valley College, where he received the college’s first Outstanding Professor Award in 1986. For many years he was also a visiting scholar at UC Riverside. Neil was the founding author of Campbell Biology.
  • Table of Contents

    Unit 1: The Chemistry of Life

    1. Evolution, the Themes of Biology, and Scientific Inquiry

    2. The Chemical Context of Life

    3. Water and Life

    4. Carbon and the Molecular Diversity of LIfe

    5. The Structure and Function of Large Biological Molecules

    Unit 2: The Cell

    1. A Tour of the Cell

    2. Membrane Structure and Function

    3. An Introduction to Metabolism

    4. Cellular Respiration and Fermentation

    5. Photosynthesis

    6. Cell Communication

    7. The Cell Cycle

    Unit 3: Genetics

    1. Meiosis and Sexual Life Cycles

    2. Mendel and the Gene Idea

    3. The Chromosomal Basis of Inheritance

    4. The Molecular Basis of Inheritance

    5. Gene Expression: from Gene to Protein

    6. Regulation of Gene Expression

    7. Viruses

    8. DNA Tools and Biotechnology

    9. Genomes and Their Evolution

    Unit 4: Mechanisms of Evolution

    1. Descent with Modification: A Darwinian View of Life

    2. The Evolution of Populations

    3. The Origin of Species

    4. The History of Life on Earth

    Unit 5: The Evolutionary History of Biological Diversity

    1. Phylogeny and the Tree of Life

    2. Bacteria and Archaea

    3. Protists

    4. Plant Diversity I: How Plants Colonized Land

    5. Plant Diversity II: The Evolution of Seed Plants

    6. Fungi

    7. An Overview of Animal Diversity

    8. An Introduction to Invertebrates

    9. The Origin and Evolution of Vertebrates

    Unit 6: Plant Form and Function

    1. Vascular Plant Structure, Growth, and Development

    2. Resource Acquisition and Transport in Vascular Plants

    3. Soil and Plant Nutrition

    4. Angiosperm Reproduction and biotechnology

    5. Plant Responses to Internal and External Signals

    Unit 7: Animal Form and Function

    1. Basic Principles of Animal Form and Function

    2. Animal Nutrition

    3. Circulation and Gas Exchange

    4. The Immune System

    5. Osmoregulation and Excretion

    6. Hormones and the Endocrine System

    7. Animal Reproduction

    8. Animal Development

    9. Neurons, Synapses, and signaling

    10. Nervous Systems

    11. Sensory and Motor Mechanisms

    12. Animal Behavior

    Unit 8: Ecology

    1. An Introduction to Ecology and the Biosphere

    2. Population Ecology

    3. Community Ecology

    4. Ecosystems and Restoration Ecology

    5. Conservation Biology and Global Change

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